K. Aizaki and Y. Yusa, Learned predator recognition in a freshwater snail, 2010.

C. N. Bedore, S. M. Kajiura, and S. Johnsen, Freezing behaviour facilitates bioelectric crypsis in cuttlefish faced with predation risk, Proc. R. Soc. B, vol.282, 1820.

J. G. Boal and D. K. Golden, Distance chemoreception in the common cuttlefish, Sepia officinalis (Mollusca, Cephalopoda), Journal of Experimental Marine Biology and Ecology, vol.235, issue.2, pp.187-194, 1999.

J. G. Boal and J. N. Ni, Ventilation rate of cuttlefish, Sepia officinalis, in response to visual stimuli, Veliger, vol.39, issue.4, pp.342-347, 1996.

M. E. Bouton, Learning and behavior: A contemporary synthesis, vol.xiii, 2007.

M. A. Sunderland, . Us:-sinauer, and . Associates,

P. R. Boyle, Cephalopod life cycles: Volume II: Comparative Reviews, 1987.

G. E. Brown and R. J. Smith, Acquired predator recognition in juvenile rainbow trout (Oncorhynchus mykiss): conditioning hatchery-reared fish to recognize chemical cues of a predator, Canadian Journal of Fisheries and Aquatic Sciences, vol.55, issue.3, pp.611-617, 1998.

M. M. Brown, N. A. Kreiter, J. T. Maple, and J. M. Sinnott, Silhouettes elicit alarm calls from captive vervet monkeys (Cercopithecus aethiops), Journal of Comparative Psychology, vol.106, issue.4, p.350, 1992.

D. P. Chivers and R. J. Smith, Chemical alarm signalling in aquatic predator-prey systems: a review and prospectus, vol.5, pp.338-352, 1998.

D. Colombelli-négrel, M. E. Hauber, and S. Kleindorfer, Prenatal learning in an Australian songbird: habituation and individual discrimination in superb fairy-wren embryos, Proceedings of the Royal Society B: Biological Sciences, vol.281, 1797.

E. Curio, U. Ernst, and W. Vieth, Cultural transmission of enemy recognition: one function of mobbing, Science, vol.202, issue.4370, pp.899-901, 1978.

S. Dalesman, S. D. Rundle, R. A. Coleman, and P. A. Cotton, Cue association and antipredator behaviour in a pulmonate snail, Lymnaea stagnalis, Animal Behaviour, vol.71, issue.4, pp.789-797, 2006.

S. Dalesman, S. D. Rundle, and P. A. Cotton, Predator regime influences innate antipredator behaviour in the freshwater gastropod Lymnaea stagnalis, Freshwater Biology, vol.52, issue.11, pp.2134-2140, 2007.

A. Darmaillacq, C. Lesimple, and L. Dickel, Embryonic visual learning in the cuttlefish, Sepia officinalis, Animal Behaviour, vol.76, issue.1, pp.131-134, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00866979

A. Darmaillacq, N. Mezrai, C. E. O'brien, and L. Dickel, Visual ecology and the development of visually guided behavior in the cuttlefish, Frontiers in Physiology, vol.8, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01542696

C. D. Derby, Cephalopod Ink: Production, Chemistry, Functions and Applications, Marine Drugs, vol.12, issue.5, pp.2700-2730, 2014.

L. M. Dill, Adaptive flexibility in the foraging behavior of fishes, Canadian Journal of Fisheries and Aquatic Sciences, vol.40, issue.4, pp.398-408, 1983.

D. L. Dixson, M. S. Pratchett, and P. L. Munday, Reef fishes innately distinguish predators based on olfactory cues associated with recent prey items rather than individual species, Animal Behaviour, vol.84, issue.1, pp.45-51, 2012.

K. J. Epp and C. R. Gabor, Innate and learned predator recognition mediated by chemical signals in Eurycea nana, Ethology, vol.114, issue.6, pp.607-615, 2008.

M. Fendt, Exposure to Urine of Canids and Felids, but not of Herbivores, Induces Defensive Behavior in Laboratory Rats, Journal of Chemical Ecology, vol.32, issue.12, 2006.

M. C. Ferrari and D. P. Chivers, Latent inhibition of predator recognition by embryonic amphibians, Biology Letters, vol.5, issue.2, pp.160-162, 2009.

M. C. Ferrari and D. P. Chivers, Sophisticated early life lessons: threat-sensitive generalization of predator recognition by embryonic amphibians, Behavioral Ecology, vol.20, issue.6, pp.1295-1298, 2009.

M. C. Ferrari and D. P. Chivers, The ghost of predation future: threat-sensitive and temporal assessment of risk by embryonic woodfrogs, Behavioral Ecology and Sociobiology, vol.64, issue.4, pp.549-555, 2010.

M. C. Ferrari, A. L. Crane, and D. P. Chivers, Certainty and the cognitive ecology of generalization of predator recognition, Animal Behaviour, vol.111, pp.207-211, 2016.

M. C. Ferrari, A. K. Manek, and D. P. Chivers, Temporal learning of predation risk by embryonic amphibians, Biology Letters, vol.6, issue.3, pp.308-310, 2010.

M. C. Ferrari, F. Messier, and D. P. Chivers, Threat-sensitive learning of predators by larval mosquitoes Culex restuans, Behavioral Ecology and Sociobiology, vol.62, issue.7, pp.1079-1083, 2008.

G. Fiorito, A. Affuso, J. Basil, A. Cole, P. De-girolamo et al., , 2015.

, Guidelines for the Care and Welfare of Cephalopods in Research-A consensus based on an initiative by CephRes, FELASA and the Boyd Group, Laboratory Animals, vol.49, issue.2_suppl, pp.1-90

T. S. Garcia, J. C. Urbina, E. M. Bredeweg, and M. C. Ferrari, Embryonic learning and developmental carry-over effects in an invasive anuran, Oecologia, vol.184, issue.3, pp.623-631, 2017.

W. F. Gilly and M. T. Lucero, Behavioral responses to chemical stimulation of the olfactory organ in the squid Loligo opalescens, Journal of Experimental Biology, vol.162, issue.1, pp.209-229, 1992.

J. L. Golub, Embryonic learning in threespine stickleback (Gasterosteus aculeatus), 2013.

R. A. Griffiths, L. Schley, P. E. Sharp, J. L. Dennis, and A. Roman, Behavioural responses of Mallorcan midwife toad tadpoles to natural and unnatural snake predators, Animal Behaviour, vol.55, issue.1, pp.207-214, 1998.

M. Guibé, N. Poirel, O. Houdé, and L. Dickel, Food imprinting and visual generalization in embryos and newly hatched cuttlefish, Sepia officinalis, Animal Behaviour, vol.84, issue.1, pp.213-217, 2012.

R. T. Hanlon and J. B. Messenger, Cephalopod behaviour, 2018.

E. J. Hartman and M. V. Abrahams, Sensory compensation and the detection of predators: the interaction between chemical and visual information, Proceedings of the Royal Society of London B: Biological Sciences, vol.267, pp.571-575, 1443.

,

L. A. Hawkins, A. E. Magurran, and J. D. Armstrong, Innate predator recognition in newly-hatched Atlantic salmon, Behaviour, issue.10, pp.1249-1262, 2004.

S. M. Hirsch and R. C. Bolles, On the Ability of Prey to Recognize Predators, Zeitschrift Für Tierpsychologie, vol.54, issue.1, pp.71-84, 1980.

I. Karplus and D. Algom, Visual Cues for Predator Face Recognition by Reef Fishes, 1981.

. Zeitschrift-für-tierpsychologie, , vol.55, pp.343-364

L. B. Kats and L. M. Dill, The scent of death: chemosensory assessment of predation risk by prey animals, vol.5, pp.361-394, 1998.

J. L. Kelley and A. E. Magurran, Learned predator recognition and antipredator responses in fishes, Fish and Fisheries, vol.4, issue.3, pp.216-226, 2003.

T. Kindermann, B. M. Siemers, and M. Fendt, Innate or learned acoustic recognition of avian predators in rodents, Journal of Experimental Biology, vol.212, issue.4, pp.506-513, 2009.

,

A. J. King and S. A. Adamo, The ventilatory, cardiac and behavioural responses of resting cuttlefish (Sepia officinalis L.) to sudden visual stimuli, Journal of Experimental Biology, vol.209, issue.6, pp.1101-1111, 2006.

M. Lee, C. Lin, C. Chiao, and C. Lu, Reproductive Behavior and Embryonic Development of the Pharaoh Cuttlefish, Sepia pharaonis (Cephalopoda: Sepiidae), p.55, 2016.

Y. Lee, A. Darmaillacq, L. Dickel, and C. Chiao, Effects of embryonic exposure to predators on the postnatal defensive behaviors of cuttlefish, Animal Behaviour
URL : https://hal.archives-ouvertes.fr/hal-02432982

M. T. Lucero, H. Farrington, and W. F. Gilly, Quantification of L-dopa and dopamine in squid ink: implications for chemoreception, The Biological Bulletin, vol.187, issue.1, pp.55-63, 1994.

F. Madaras, J. P. Gerber, F. Peddie, and M. J. Kokkinn, The effect of sampling methods on the apparent constituents of ink from the squid Sepioteuthis australis, Journal of Chemical Ecology, vol.36, issue.11, pp.1171-1179, 2010.

A. Mathis, M. C. Ferrari, N. Windel, F. Messier, and D. P. Chivers, Learning by embryos and the ghost of predation future, Proceedings of the Royal Society B: Biological Sciences, vol.275, pp.2603-2607, 1651.

A. Mathis and R. J. Smith, Fathead minnows, Pimephales promelas, learn to recognize northern pike, Esox lucius, as predators on the basis of chemical stimuli from minnows in the pike's diet, Animal Behaviour, vol.46, issue.4, pp.645-656, 1993.

,

N. Mezrai, C. Chiao, L. Dickel, and A. Darmaillacq, Different timings for the onset of visual and chemosensory systems during embryonic development in two closely related cuttlefish species

Á. Miklósi, P. Pongrácz, and V. Csányi, The ontogeny of antipredator behavior in paradise fish larvae (Macropodus opercularis) IV. The effect of exposure to siblings, Developmental Psychobiology, vol.30, issue.4, pp.283-291, 1997.

R. S. Mirza, M. C. Ferrari, J. M. Kiesecker, and D. P. Chivers, Responses of American toad tadpoles to predation cues: behavioural response thresholds, threatsensitivity and acquired predation recognition, Behaviour, vol.143, issue.7, pp.877-889, 2006.

,

R. Misslin, The defense system of fear: behavior and neurocircuitry, Neurophysiologie Clinique/Clinical Neurophysiology, vol.33, issue.2, pp.9-10, 2003.

C. E. O'brien, N. Mezrai, A. Darmaillacq, and L. Dickel, Behavioral development in embryonic and early juvenile cuttlefish (Sepia officinalis), Developmental Psychobiology, vol.59, issue.2, pp.145-160, 2017.

L. J. Oulton, V. Haviland, and C. Brown, Predator Recognition in Rainbowfish, Melanotaenia duboulayi, Embryos. PLOS ONE, vol.8, issue.10, p.76061, 2013.

,

S. W. Porges and D. C. Raskin, Respiratory and heart rate components of attention, Journal of Experimental Psychology, vol.81, issue.3, p.497, 1969.

G. Prota, J. P. Ortonne, C. Voulot, C. Khatchadourian, G. Nardi et al., , 1981.

, Occurrence and properties of tyrosinase in the ejected ink of cephalopods, Comparative Biochemistry and Physiology Part B: Comparative Biochemistry, vol.68, issue.3, pp.415-419

J. E. Richards and B. J. Casey, Heart rate variability during attention phases in young infants, Psychophysiology, vol.28, issue.1, pp.43-53, 1991.

R. Rochette, D. J. Arsenault, B. Justome, and J. H. Himmelman, Chemically-mediated predator-recognition learning in a marine gastropod, Écoscience, vol.5, issue.3, pp.353-360, 1998.

,

S. Romagny, A. Darmaillacq, M. Guibé, C. Bellanger, and L. Dickel, Feel, smell and see in an egg: emergence of perception and learning in an immature invertebrate, the cuttlefish embryo, The Journal of Experimental Biology, vol.215, issue.23, pp.4125-4130, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00862809

P. Saglio and A. Mandrillon, Embryonic experience to predation risk affects tadpoles of the common frog (Rana temporaria), Archiv Für Hydrobiologie, vol.166, issue.4, pp.505-523, 2006.
URL : https://hal.archives-ouvertes.fr/hal-01453730

S. P. Saunders, T. W. Ong, and F. J. Cuthbert, Auditory and visual threat recognition in captive-reared Great Lakes piping plovers (Charadrius melodus), Applied Animal Behaviour Science, vol.144, issue.3, pp.153-162, 2013.

A. C. Utne-palm, Response of naive two-spotted gobies Gobiusculus flavescens to visual and chemical stimuli of their natural predator, cod Gadus morhua, Marine Ecology Progress Series, vol.218, pp.267-274, 2001.

T. Vehanen, Adaptive flexibility in the behaviour of juvenile Atlantic salmon: shortterm responses to food availability and threat from predation, Journal of Fish Biology, vol.63, issue.4, pp.1034-1045, 2003.

M. J. Wells, Sensitization and the Evolution of Associative Learning, Neurobiology of Invertebrates, pp.391-411, 1968.

B. D. Wisenden, Chemically mediated strategies to counter predation, Sensory processing in aquatic environments, pp.236-251, 2003.

B. D. Wisenden, D. P. Chivers, and R. J. Smith, Learned Recognition of Predation Risk by Enallagma Damselfly Larvae (Odonata, Zygoptera) on the Basis of Chemical Cues, Journal of Chemical Ecology, vol.23, issue.1, pp.137-151, 1997.

,

B. D. Wisenden and M. C. Millard, Aquatic flatworms use chemical cues from injured conspecifics to assess predation risk and to associate risk with novel cues, Animal Behaviour, vol.62, issue.4, pp.761-766, 2001.

D. R. Woody and A. Mathis, Acquired recognition of chemical stimuli from an unfamiliar predator: associative learning by adult newts, Notophthalmus viridescens, Copeia, issue.4, pp.1027-1031, 1998.

*. Wright, T. F. Eberhard, *. , J. R. Hobson, E. A. Avery et al., , 2010.

, Behavioral flexibility and species invasions: the adaptive flexibility hypothesis, Ethology Ecology & Evolution, vol.22, issue.4, pp.393-404

,

J. Z. Young, Learning and Discrimination in the Octopus, Biological Reviews, vol.36, issue.1, pp.32-95, 1961.