R. Amara, P. Laffargue, J. M. Dewarumez, C. Maryniak, F. Lagardère et al., Feeding ecology and growth of O-group flatfish (sole, dab and plaice) on a nursery ground (Southern Bight of the North Sea), J. Fish Biol, vol.58, pp.788-803, 2001.

S. C. Anderson, Improving estimates of population status and trend with superensemble models. Fish Fish, vol.18, pp.732-741, 2017.

H. Asmus, Secondary production of an intertidal mussel bed community related to its storage and turnover compartments, Mar. Ecol. Prog. Ser, vol.39, p.39251, 1987.

D. Baird and R. E. Ulanowicz, The seasonal dynamics of the Chesapeake Bay ecosystem, Ecol. Monogr, vol.59, pp.329-364, 1989.

A. C. Benke and A. D. Huryn, Benthic invertebrate production-facilitating answers to ecological riddles in freshwater ecosystems, J. North Am. Benthol. Soc, vol.29, pp.264-285, 2010.

J. J. Beukema, Seasonal changes in the biomass of the macro-benthos of a tidal flat area in the Dutch Wadden Sea, Neth. J. Sea Res, vol.8, pp.90028-90031, 1974.

J. J. Beukema and W. De-bruin, Seasonal changes in dry weight and chemical composition of the soft parts of the tellinid bivalve Macoma balthica in the Dutch Wadden Sea, Neth. J. Sea Res, vol.11, pp.90020-90025, 1977.

J. J. Beukema and R. Dekker, Evaluation of Brey's production/biomass model on the basis of a long-term data set on a clam population, Mar. Ecol. Prog. Ser, vol.489, pp.163-175, 2013.

J. J. Beukema, G. C. Cadée, R. Dekker, and C. J. Philippart, Annual and spatial variability in gains of body weight in Macoma balthica (L.): Relationships with food supply and water temperature, J. Exp. Mar. Biol. Ecol, vol.457, pp.105-112, 2014.

S. G. Bolam, C. R. Barrio-frojan, and J. D. Eggleton, Macrofaunal production along the UK continental shelf, J. Sea Res, vol.64, pp.166-179, 2010.

S. G. Bolam and J. D. Eggleton, Macrofaunal production and biological traits: Spatial relationships along the UK continental shelf, J. Sea Res, vol.88, pp.47-58, 2014.

D. G. Boyce, M. R. Lewis, and B. Worm, Global phytoplankton decline over the past century, Nature, vol.466, pp.591-596, 2010.

T. Brey, Estimating productivity of macrobenthic invertebrates from biomass and mean individual weight, Meeresforsch, vol.32, pp.329-343, 1990.

T. Brey, Population dynamics in benthic invertebrates. A virtual handbook. Version 01.2, 2001.

T. Brey, A multi-parameter artificial neural network model to estimate macrobenthic invertebrate productivity and production: Macrobenthic production model, Limnol. Oceanogr.: Methods, vol.10, pp.581-589, 2012.

T. Brey, A. Jarre-teichmann, and O. Borlich, Artificial neural network versus multiple linear regression: Predicting P/B ratios from empirical data, Mar. Ecol. Prog. Ser, vol.140, pp.251-256, 1996.

T. Brey, C. Müller-wiegmann, Z. M. Zittier, and W. Hagen, Body composition in aquatic organisms -a global data bank of relationships between mass, elemental composition and energy content, J. Sea Res, vol.64, pp.334-340, 2010.

L. Bunting, P. R. Leavitt, G. L. Simpson, B. Wissel, K. R. Laird et al., Increased variability and sudden ecosystem state change in Lake Winnipeg, Canada, caused by 20th century agriculture, Limnol. Oceanogr, vol.61, pp.2090-2107, 2016.

B. J. Burd, T. A. Macdonald, and A. Van-roodselaar, Towards predicting basin-wide invertebrate organic biomass and production in marine sediments from a coastal sea, PLoS One, vol.7, p.40295, 2012.

R. Callaway, Tube worms promote community change, Mar. Ecol. Prog. Ser, vol.308, pp.49-60, 2006.

R. Callaway, N. Desroy, S. F. Dubois, J. Fournier, M. Frost et al., Ephemeral bioengineers or reef-building polychaetes: How stable are aggregations of the tube worm Lanice conchilega (Pallas, 1766)?, Integr. Comp. Biol, vol.50, pp.237-250, 2010.
URL : https://hal.archives-ouvertes.fr/hal-00617494

J. E. Cartes, T. Brey, J. C. Sorbe, and F. Maynou, Comparing production biomass ratios of benthos and suprabenthos in macrofaunal marine crustaceans, Can. J. Fish. Aquat. Sci, vol.59, pp.1616-1625, 2002.

R. Coma, M. Ribes, J. Gili, and M. Zabala, Seasonality in coastal benthic ecosystems, Trends Ecol. Evol, vol.15, pp.448-453, 2000.

C. Conq, J. J.-c.-dauvin, and . Lorgeré, Température et salinité de l'eau de mer au large de Roscoff de, Cah. Biol. Mar, vol.39, pp.207-212, 1991.

C. J. Curtis and G. L. Simpson, Trends in bulk deposition of acidity in the UK, 1988-2007, assessed using additive models, Ecol. Indic, vol.37, pp.274-286, 2014.

R. M. Cushman, H. H. Shugart, S. G. Hildebrand, and J. W. Elwood, The effect of growth curve and sampling regime on instantaneous-growth, removal-summation, and Hynes/Hamilton estimates of aquatic insect production: A computer simulation, Limnol. Oceanogr, vol.23, pp.184-189, 1978.

M. Cusson and E. Bourget, Global patterns of macroinvertebrate production in marine benthic habitats, Mar. Ecol. Prog. Ser, vol.297, pp.1-14, 2005.

M. Cusson, J. Plante, and C. Genest, Effect of different sampling designs and methods on the estimation of secondary production: A simulation, Limnol. Oceanogr.: Methods, vol.4, pp.38-48, 2006.

J. Dauvin, The fine sand Abra alba community of the Bay of Morlaix twenty years after the Amoco Cadiz oil spill, Mar. Pollut. Bull, vol.36, pp.58-65, 1998.

J. Dauvin, The muddy fine sand Abra alba-Melinna palmata community of the Bay of Morlaix twenty years after the Amoco Cadiz oil spill, Mar. Pollut. Bull, vol.40, pp.528-536, 2000.

J. Dauvin, M. Joncourt, D. Latrouite, and G. Raguénès, Température et salinité de l'eau de mer au large de Roscoff de, Cah. Biol. Mar, vol.30, pp.5-10, 1983.

D. Smet and B. , Biogenic reefs affect multiple components of intertidal soft-bottom benthic assemblages: The Lanice conchilega case study, Estuar. Coast. Shelf Sci, vol.152, pp.44-55, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01087288

R. Degen, L. L. Jørgensen, P. Ljubin, I. H. Ellingsen, H. Pehlke et al., Patterns and drivers of megabenthic secondary production on the Barents Sea shelf, Mar. Ecol. Prog. Ser, vol.546, pp.1-16, 2016.

N. Desroy and C. Retière, Long-term changes in muddy fine sand community of the Rance Basin: Role of recruitment, J. Mar. Biol. Assoc. U.K, vol.81, pp.553-564, 2001.

J. M. Dewarumez, A. Belgrano, J. Craeymeersch, S. Duquesne, C. Heip et al., Influence de la circulation des masses d'eaux dans la dynamique du peuplement Abra alba de la Baie sud de la Mer du Nord, J. Rech. Oceanogr, vol.18, pp.1-4, 1993.

M. Dolbeth, A. I. Lillebø, P. G. Cardoso, S. M. Ferreira, and M. A. , Annual production of estuarine fauna in different environmental conditions: An evaluation of the estimation methods, J. Exp. Mar. Biol. Ecol, vol.326, pp.115-127, 2005.

M. Dolbeth, M. Cusson, R. Sousa, M. A. Pardal, and Y. T. Prairie, Secondary production as a tool for better understanding of aquatic ecosystems, Can. J. Fish. Aquat. Sci, vol.69, pp.1230-1253, 2012.

I. C. Field, C. J. Bradshaw, H. R. Burton, and M. A. Hindell, Differential resource allocation strategies in juvenile elephant seals in the highly seasonal Southern Ocean, Mar. Ecol. Prog. Ser, vol.331, pp.281-290, 2007.

C. L. Frid, J. B. Buchanan, and P. R. Garwood, Variability and stability in benthos: Twenty-two years of monitoring off Northumberland, ICES J. Mar. Sci, vol.53, pp.978-980, 1996.

J. M. Fromentin and F. Ibanez, Year-to-year changes in meteorological features of the French coast area during the last half-century-examples of 2 biological responses, Oceanol. Acta, vol.17, pp.285-296, 1994.

J. M. Fromentin, F. Ibanez, J. Dauvin, J. M. Dewarumez, and B. Elkaim, Long-term changes of four macrobenthic assemblages from 1978 to 1992, J. Mar. Biol. Assoc. U. K, vol.77, pp.287-310, 1997.

M. M. Fuhrmann, T. Pedersen, V. Ramasco, and E. M. Nilssen, Macrobenthic biomass and production in a heterogenic subarctic fjord after invasion by the red king crab, J. Sea Res, vol.106, pp.1-13, 2015.

K. Ghertsos, C. Luczak, J. Dewarumez, and J. Dauvin, Influence of spatial scales of observation on temporal change in diversity and trophic structure of fine-sand communities from the English Channel and the southern North Sea, ICES J. Mar. Sci, vol.57, pp.1481-1487, 2000.

P. A. Henderson, R. M. Seaby, and J. R. Somes, A 25-year study of climatic and density-dependent population regulation of common shrimp Crangon crangon (Crustacea: Caridea) in the Bristol Channel, J. Mar. Biol. Assoc. U. K, vol.86, pp.287-298, 2006.

R. R. Hessler and H. L. Sanders, Faunal diversity in the deep-sea. Deep-Sea Res, Oceanogr. Abstr, vol.14, pp.65-78, 1967.

A. S. Jung, R. Dekker, M. Germain, C. J. Philippart, J. I. Witte et al., Long-term shifts in intertidal predator and prey communities in the Wadden Sea and consequences for food requirements and supply, Mar. Ecol. Prog. Ser, vol.579, pp.37-53, 2017.

M. K?dra, P. E. Renaud, H. Andrade, I. Goszczko, and W. G. Ambrose, Benthic community structure, diversity, and productivity in the shallow Barents Sea bank (Svalbard Bank), Mar. Biol, vol.160, pp.805-819, 2013.

B. R. Kuipers, P. D. Wilde, and F. Creutzberg, Energy flow in a tidal flat ecosystem, Mar. Ecol. Prog. Ser, vol.5, pp.215-221, 1981.

C. Luczak, Dynamique du recrutement au sein du peuplement à Abra alba dans la baie sud de la Mer du Nord: Approche systémique, 1996.

C. Luczak, J. Dewarumez, and K. Essink, First record of the American jack knife clam Ensis directus on the French coast of the North Sea, J. Mar. Biol. Assoc. U. K, vol.73, pp.233-235, 1993.

A. Morin, T. A. Mousseau, and D. A. Roff, Accuracy and precision of secondary production estimates, Limnol. Oceanogr, vol.32, pp.1342-1352, 1987.

A. Morin and N. Bourassa, Modèles empiriques de la production annuelle et du rapport P/B d'invertébrés benthiques d'eau courante, Can. J. Fish. Aquat. Sci, vol.49, pp.532-539, 1992.

A. P. Mucha, M. T. Vasconcelos, and A. A. Bordalo, Spatial and seasonal variations of the macrobenthic community and metal contamination in the Douro estuary (Portugal), Mar. Environ. Res, vol.60, pp.531-550, 2005.

K. Ono, The importance of length and age composition data in statistical age-structured models for marine species, ICES J. Mar. Sci, vol.72, pp.31-43, 2015.

G. Passuni, Seasonality in marine ecosystems: Peruvian seabirds, anchovy, and oceanographic conditions, Ecology, vol.97, pp.182-193, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01298551

J. Paw?owska, M. W?odarska-kowalczuk, M. Zaj?czkowski, H. Nygard, and J. Berge, Seasonal variability of meio-and macrobenthic standing stocks and diversity in an Arctic fjord (Adventfjorden, Spitsbergen), Polar Biol, vol.34, pp.833-845, 2011.

M. Petracco, R. S. Cardoso, T. N. Corbisier, and A. Turra, Secondary production of sandy beach macrofauna: An evaluation of predictive models, Estuar. Coast. Shelf Sci, vol.115, pp.359-365, 2012.

C. J. Philippart, H. M. Van-aken, J. J. Beukema, O. G. Bos, G. C. Cadée et al., Climate-related changes in recruitment of the bivalve Macoma balthica, Limnol. Oceanogr, vol.48, pp.2171-2185, 2003.

L. Pihl, Food selection and consumption of mobile epibenthic fauna in shallow marine areas, Mar. Ecol. Prog. Ser, vol.22, pp.169-179, 1985.

C. Plante and J. A. Downing, Production of freshwater invertebrate populations in lakes, Can. J. Fish. Aquat. Sci, vol.46, pp.1489-1498, 1989.

. R-core-team, R: A language and environment for statistical computing. R Foundation for Statistical Computing, 2017.

H. Reiss and I. Kröncke, Seasonal variability of infaunal community structures in three areas of the North Sea under different environmental conditions, Estuar. Coast. Shelf Sci, vol.65, pp.253-274, 2005.

H. Reiss, Effects of fishing disturbance on benthic communities and secondary production within an intensively fished area, Mar. Ecol. Prog. Ser, vol.394, pp.201-213, 2009.

A. I. Robertson, The relationship between annual production: Biomass ratios and lifespans for marine macrobenthos, Oecologia, vol.38, pp.193-202, 1979.

T. A. Schlacher and T. H. Wooldridge, How sieve mesh size affects sample estimates of estuarine benthic macrofauna, J. Exp. Mar. Biol. Ecol, vol.201, issue.95, pp.198-198, 1996.

D. S. Schoeman, M. Wheeler, and M. Wait, The relative accuracy of standard estimators for macrofaunal abundance and species richness derived from selected intertidal transect designs used to sample exposed sandy beaches, Estuar. Coast. Shelf Sci, vol.58, pp.38-44, 2003.

P. Schwinghamer, B. Hargrave, D. Peer, and C. M. Hawkins, Partitioning of production and respiration among size groups of organisms in an intertidal benthic community, Mar. Ecol. Prog. Ser, vol.31, pp.131-142, 1986.

M. Sprung, Estimating macrobenthic secondary production from body weight and biomass: A field test in a non-boreal intertidal habitat, Mar. Ecol. Prog. Ser, vol.100, pp.103-109, 1993.

A. Tableau, H. L. Bris, and A. Brind'amour, Available benthic energy coefficient (ABEC): A generic tool to estimate the food profitability in coastal fish nurseries, Mar. Ecol. Prog. Ser, vol.522, pp.203-218, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01453882

D. W. Townsend, M. D. Keller, M. E. Sieracki, and S. G. , Ackleson. 1992. Spring phytoplankton blooms in the absence of vertical water column stratification, Nature, vol.360, pp.59-62

M. L. Tumbiolo and J. A. Downing, An empirical model for the prediction of secondary production in marine benthic invertebrate populations, Mar. Ecol. Prog. Ser, vol.114, pp.165-174, 1994.

R. W. Virnstein, The importance of predation by crabs and fishes on benthic infauna in Chesapeake Bay, Ecology, vol.58, pp.1199-1217, 1977.

B. A. Walther and J. L. Moore, The concepts of bias, precision and accuracy, and their use in testing the performance of species richness estimators, with a literature review of estimator performance, Ecography, vol.28, pp.815-829, 2005.

G. Walther, E. Post, P. Convey, A. Menzel, C. Parmesan et al., Ecological responses to recent climate change, Nature, vol.416, pp.389-395, 2002.

K. H. Wiltshire, A. M. Malzahn, K. Wirtz, W. Greve, S. Janisch et al., Resilience of North Sea phytoplankton spring bloom dynamics: An analysis of long-term data at Helgoland Roads, Limnol. Oceanogr, vol.53, pp.1294-1302, 2008.

D. Woehrling, A. Lefebvre, G. Le-fèvre-lehoërff, and R. Delesmont, Seasonal and longer term trends in sea temperature along the French North Sea coast, J. Mar. Biol. Assoc. U. K, vol.85, pp.39-48, 1975.

M. C. Wong, C. H. Peterson, and M. F. Piehler, Evaluating estuarine habitats using secondary production as a proxy for food web support, Mar. Ecol. Prog. Ser, vol.440, pp.11-25, 2011.

S. N. Wood, Generalized additive models: An introduction with R, 2006.

S. N. Wood, Fast stable restricted maximum likelihood and marginal likelihood estimation of semiparametric generalized linear models, J. R. Stat. Soc. Ser. B Stat. Methodol, vol.73, pp.3-36, 2011.

R. Zühlke, Polychaete tubes create ephemeral community patterns: Lanice conchilega (Pallas, 1766) associations studied over six years, J. Sea Res, vol.46, pp.91-91, 2001.

L. Zwarts and J. H. Wanink, How the food supply harvestable by waders in the Wadden Sea depends on the variation in energy density, body weight, biomass, burying depth and behaviour of tidal-flat invertebrates, Neth. J. Sea Res, vol.31, issue.93, pp.90059-90061, 1993.